In their Rezzed debut, Playtonic confirmed that along with having different characters be the Heroes for different games in their universe, so too would the 'Villains' be different. It sounds like they're trying to make a realistic series of situations, whereby characters moralities are fluid depending on who they interact with. I imagine there will be one overarching villain, at least for the first game, but other situations depend on the other characters - perhaps a group of Good and a group of Bad, with different 'Villains' holding the power depending on the game, or perhaps a completely fluid morality where, say, a Beaver is the enemy of an Otter in one game, and in another the Otter is the enemy of a jungle cat - we'll just have to wait and see. From the silhouettes shown at their Rezzed debut, they appear to be including animal and human characters, perhaps also robotic or humanoid creatures. Very eager to see what their minds come up with!
Came to save my sister, stayed for the quiz show.